- Project of the MAST and the INCO programme of the European Union -


TOP 

 
 

- 10. ISSUE (March 1999) -

 

. 
BASYS - Baltic Sea System Study  
 

EU Research Project



 
CONTENTS
 

[Back to Contents]


 
EDITORIAL NOTE
  The first three scientific papers in the present NEWSLETTER (Vuorinen et al, Kostrychkina et al., Kornilovs) are coming from subproject 1B (Historical data) and are devoted to the changes in plankton and fish on climatic scales. The response on natural fluctuations in salinity (atmosphrically-driven inflow events) and man-made variations in nutrients (eutrophication effects) are analysed. Their causal relation to diversity and abundance as well as interaction schemes is discussed.
    The fourth contribution (Lehmann et al.) presents mean circulation patterns of the Baltic Sea, which are stable characteristics of the system. Transport and mixing features turn out to be regionally specific and shall be adequately assembled for basin-wide investigations.
 
[Back to Contents]



 
MANAGEMENT NOTES

Current state of the project

   To promote the information exchange during last phase of the project we made available some information: (Christoph Zülicke, IO Warnemünde, February 1999, mailto: christoph.zuelicke@io-warnemuende.de)
 
[Back to Contents]

5th meeting of SP6 (Baltic Sea Ice)

Date / Place: Juliusruh, Rügen (Germany), 28 May 1998

Participants:

Lars Axell, SMHI, Sweden
Jari Haapala (chairman), UH, Finland
Andreas Lehmann (rapporteur), IfM­Kiel, Germany
Anders Omstedt, SMHI, Sweden
Markus Meier, Rossby Center, Sweden (guest)

1. General

The 5th coordination meeting of BASYS/SP6 was set up in order to prepare the 24­month management report and the 2nd scientific annual progress report. The meeting took place during the 2nd Study Conference on BALTEX, Hotel Aquamaris, Rügen, Germany, 25­29 May 1998.
    Since most of the participants reported on the progress of their scientific work in connection with BASYS/SP6 during the BALTEX Study Conference (BALTEX, 1998), no additional scientific reports were given during the coordination meeting.

2. BALTEX SSG­Meeting, 27 May 1998

Anders Omstedt reported briefly on the BALTEX SSG­Meeting, which took place the day before. The SSG together with the BALTEX Task Force group are concentrating on the Main BALTEX Experiment (BRIDGE, Strategic Plan, Int. BALTEX Secetariat, Pub. No. 9) which is scheduled to take place from October 1999 to December 2001. Two papers have been prepared by the Task Force and the BALTEX Secretariat. The first paper is a Technical Implementation Plan (TIP) for BRIDGE and the second one is the Interim Memorandum of Understanding for the Conduct of BRIDGE (IMOU). The purpose of the second paper is to summarize the BRIDGE strategy as given in the Strategic Plan and to indicate the distribution of activities and responsibilities among parties for the preparation and conduct of BRIDGE. It is thereby understood that the realisation of both the preparation and conduct of BRIDGE activities will depend on satisfactory solutions of financial, logistical and manpower problems. However, while the parties have confirm intention to adhere to the terms of the IMOU, it is understood that the document has no legal implications. To contribute to BRIDGE, the different institutes should give a commitment to the IMOU which kind of tasks they are able to perform.

3. BASYS annual scientific report

Andreas Lehmann informed the group about the last meeting of the scientific steering committee (8 January 1998, Copenhagen, Denmark). The minutes of the meeting as well as other important informations may be found at the BASYS homepage:
http://www.io-warnemuende.de/Projects/Basys.
The next annual progress report together with the 24­month managment report will be delivered to MAST in August 1998. How the reports should be structure can also be found at the BASYS homepage. The subproject overview will be written by the subproject coordinator (W. Krauss, IfM­Kiel). This will be a synthesis of the scientific contributions of the members of the subproject. The first version of the reports has to be sent to the project coordinator until 15 July 1998. The next BASYS annual conference will take place in Stockholm, 23­25 September 1998.

4. Subproject Overview

Task 6.5­6.6 Model runs for 3 winters
    Andreas Lehmann reported briefly that the 3 winters (1986/87, 1992/93 and 1993/94) have already been simulated with the coupled sea ice­ocean model. However, verification and analysis of the simulations with corresponding observations and with results from the PROBE­Baltic and the Finnish coupled ice­ocean model has not been performed yet. The progress in 6.5­6.6 will be summarized by the IfM­Kiel. Tasks 6.5­6.6 are on schedule.

Task 6.7 Critical validation of the Baltic Sea model with selected observational data sets
    Andreas Lehmann pointed out that this task is partly covered by 6.5­6.6, so that the results of the analysis will be a part of this task. Anders Omstedt will also contribute to results from the PROBE­Baltic. The progress will be summarized by the SMHI and theIFM. The task is on schedule.

Task 6.8 Development and comparison of vertical diffusion paramterization schemes for ice covered seas
    Since Markus Meier has officially left the subproject, his tasks have to be distributed among the members of the subproject. The participants discussed how to proceed. Markus Meier stated that he will take part in further meetings of the subproject to foster the information exchange between the group and the newly formed Rossby Center (Norrk¨oping, Sweden). The group agreed that the k - epsilon Kiel model and the Helsinki moderate resolution model Since this work has already started in the last year, the progress achieved so far will be summarized by Jari Haapala (UH) and Andreas Lehmann (IfM). From 1st of August to 31st October 1998 Jari Haapala will work in the Theoretical Department of Oceanography at the IfM­Kiel. It is planned that during his stay much of the comparison work will be done in Kiel.

Task 6.10 Examination of pack ice mechanics paramterization schemes
    At the BALTEX conference, Jari Haapala reported on modelling of the ice thickness redistribution. An extended ice classification improves the description of minimum ice strength and thus gives more realistic ice velocities when thin ice is present. Also the different thermodynamic growth/melting rates of the ice types can be introduced into the model, hence giving more detailed seasonal evolution of the pack ice. Jari Haapala and Matti Lepp¨aranta (UH) will summarize the progress of this task.

Task 6.11 Examination the role of ice melting processes in the full model
    Jari Haapala pointed out that at the UH much progress has been achieved by a study on snow and snow ice in sea ice thermodynamic modelling presented by Tuomo Saloranta (T. Saloranta, 1998). In additions, UH has performed optical sea ice measurements in the Gulf of Finland during spring, 1998. The progress of this task will be summarized by Jari Haapala with an additional contribution by Anders Omstedt. It seems that this task cannot be finalized as required (s. Technical Annex), so that the work will be continued in the 3rd year.

5. Miscellaneous

The SMHI­Meteorological data base which provides most of the atmospheric forcing data to drive the coupled sea ice­ocean models should be referenced as: SMHI­Meteorological Data Base (Lars Meuller, pers. comm.).
    A special workshop to foster the scientific synthesis of BASYS/SP6 is planned to take place in Hailuoto/Icebreaker, Finland in 15­17 March 1999.
    The next meeting of BASYS/SP6 will be in Stockholm during the 2nd BASYS annual science conference

References

    The Main BALTEX Experiment 1999­2001 ­ BRIDGE. Strategic Plan. October 1997. Int. BALTEX Secretariat, Pub. No. 9, 78 pp.
    Saloranta, T.M., 1998: Snow and snow ice in sea ice thermodynamic modelling. Report Series in Geophysics, No. 39, University of Helsinki, 84pp.
    Second Study Conference on BALTEX, Juliusruh, Island of Rügen, Germany, 25­29 May 1998. Conference Proceedings. Editors: E. Raschke and H.­J. Isemer. Int. BALTEX Secretariat, Pub. No. 11, 251 pp. May 1998, 251pp.
 
(Andreas Lehmann, Kiel, June 1998)
[Back to Contents]


SCIENTIFIC NOTES


SCIENTIFIC PAPERS

Change of plankton diversity explained by climatic factors - An Archipelago Sea case study -

Ilppo Vuorinen and Jari Hänninen
Archipelago Research Institute
University of Turku, Finland

Plankton diversity changed since 1976-77

A decrease has been documented in the abundance of neritic large Copepoda in the Baltic proper and Gulf of Finland by several authors (Lumberg & Ojaveer, 1991; Flinkman et al., 1998; Ojaveer et al., 1998 and  Vuorinen et al., 1998).  This significant decrease of zooplankton started in 1976-77, and is comparable to the ones observed by Aebisher et al. (1992) in the North Sea and Roemmich & McGowan (1995)  in California. The North Sea zooplankton decline is connected to the Gulf Stream position (Hays et al., 1993) and, ultimately to North Atlantic weather patterns (Aebischer et al., 1992), we have been looking for evidence of a climatic pacemaker also behind the baltic Sea zooplankton.

Salinity is the decisive environmental factor in the Baltic Sea

The correlation between zooplankton and salinity in the Baltic Sea is throughly demonstrated (e.g. Segerstråle, 1969; Vuorinen & Ranta, 1987; Ranta & Vuorinen, 1990; Viitasalo et al., 1995).  In practice all the  zooplankton species in the Baltic Sea react  very clearly to changes in salinity either positively or negatively , e.g Centropages hamatus adults were steadily decreasing since 1976-77, and not found at all in our samples in 1991-92, but appeared in great numbers in 1993 after the first pulse of saline water since the beginning of the stagnation period (Fig. 1).
Fig. 1: From the top down:  The river runoff to the Baltic Proper (km3, inverted, data from Bergström & Carsson 1994),  salinity changes at the Gothland Deep from 200 m depth (%o, 6 months moving average, data provided by HELCOM) and Päiväluoto, Finland from 20 m depth (%o, 12 months moving average, data provided by FIMR) and abundance of the copepod Centropages hamatus at the Island of Seili.

Furthermore the control of salinity has been shown to occur by river runoff (Launiainen & Vihma, 1990) and other weather related factors (Matthaus & Schinke, 1994).  Salinity changes of the Baltic Sea have been attributed to irregular pulses of  North Sea water penetrating the Danish Sounds. However, the occurrence of major pulses has proved unpredictable, and they were almost totally lacking during the 1980´s.  Furthermore, the observation series collected on the occurrence of  major pulses of saline water has not allowed for building up a predictive model.

Relation to weather patterns

We used transfer functions in order to model the relationship between westerly winds measured over the British Isles and the freswater runoff  to the Baltic Proper catchment area (Hänninen et al., 1999). Furthermore,  we modelled the relationship between the  NAO index and the freshwater runoff from the total Baltic Sea catchment area, and also the runoff  from the Baltic Proper cathment area.  We  also modelled the relationship between the NAO and the Baltic seawater salinity.  The westerly winds had a direct effect in the runoff of the Baltic Proper with a lag of  less than one month. The freshwater runoff responds to changes in the NAO-index with a lag of about 1-4 months depending on the runoff area in question (Fig. 2).
Fig. 2  Two years moving average of the freshwater runoff to the Baltic Sea (thick grey line, km3 month-1) and the Northern Atlantic Oscillation index (thin black line).  At the abscissa months are running from 1921 to 1991, at the left ordinate the river runoff .

Moreover, the salinities respond to changes in the runoff from the total catchment area of the Baltic with a lag of about one year.  During the years of high NAO there has been a substantial increase in the rainfall over an area extending from the Northern Germany and the British Isles over Scandinavia.  Simultaneously the Baltic Sea has experienced an almost total lack of saline water pulses which would have increased the salinity.  Thus the predictive ability of our models is based on the time lag between the cyclones forming in the Atlantic and their migration and subsequent rainfall and runoff over the Scandinavia, here exemplified with the Baltic Sea cathment area.

Possible impact on fishery

While predicting salinity changes on the basis of runoff and NAO is fairly straightforward, we feel it intriguing to have a possibility to predict more complicated ecological interactions.  Starting from the NAO, we foresee it possible to extend our prediction to cover also biological interactions, which generally seem to follow the same reasoning, although the controlling factors are more numerous (and may be working in concert or antagonistically). Our previous studies have shown the connection between seawater salinity and mesozooplankton in the Baltic Sea and, subsequently between zooplankton and it´s main predator, the Baltic herring .  Decreasing salinites during the 1980´s have brought a change in zooplankton species composition and abundance by favouring smaller mesozooplankton, which generally originates from freshwater. Simultaneously, the food composition of the Baltic herring has changed accordingly, and the herring is suffering of decrased quality and quantity of it´s staple food, the planktonic copepods (Lumberg & Ojaveer, 1991; Flinkman et al., 1998).  Thus the poor condition and starvation of  the Baltic herring as also the decline in the Baltic cod stock both occurred during the 1980´s  and  caused problems for fisheries and fisheries management are explained, and at least partly attributable to salinity changes, and ultimately to meterological control through the NAO.  While this cascade closely resembles that described by Aebischer et al. (1990), we want to point out that also the cascading trophic interaction  due to selective feeding of the herring would work in the same direction.   The increased predation by the Baltic herring which has been documented for example by Flinkman et al. (1998) has also been interpreted in the light of salinity changes: the decrease of salinity has caused a decline in the Baltic cod stocks. With decreasing cod predation the herring stocks flourish and may deplete the large zooplankton.  Even salinity as such might have an impact on herring (as well as zooplankton) directly through the osmoregulation.  Ultimately salinity-driven phytoplankton changes, which were not studied here may exert some effect up the food chain.  However, even these mechanisms having some effect to the overall plankton abundance the ultimate controlling factors are the weather patterns and the climate.

Acknowledgements

This work  has been formed in co-operation with several persons, first the herring biology research group at Archipelago Research Institute (ARI), which is lead by Dr. Marjut Rajasilta, and also research group of zooplankton ecology at the University of Helsinki lead by Dr. Markku Viitasalo, furthermore researchers at the Finnish Institute of Marine Research are acknowledged for their contribution to these ideas.

References

    Aebischer, N.J., Coulson, J.C. & Colebrook, J.M., 1990: Parallel long-term trends across four marine trophic levels and weather. -Nature 347:753-755.
    Bergström, S. & Carlsson, B., 1994: Hydrology of the Baltic Basin. Inflow of  fresh water from rivers and land for the period 1950-1990. SMHI Reports Hydrology. 7.
    Flinkman, J., Aro, E. & Vuorinen, I. & Viitasalo, M., 1998: Changes in northern Baltic zooplankton and herring nutrition from 1980s to  1990s - top-down and bottom-up processes at work. - Mar. Ecol.  Prog. Ser. 165:127-136.
    Hays, G.C., Carr, M.R. & Taylor, A.H., 1993: The Relationship between Gulf Stream position and copepod abundance derived from the Continuous Plankton Recorder Survey: separating biological signal from sampling noise. -J. Plankton Res. 15:1359-1373.
    Hänninen, J., Vuorinen, I. & Hjelt, P., 1999: Predicting oceanographical changes of the Baltic Sea on the basis of climatic factors.- a manuscript suibmitted
    Launiainen, J. & Vihma, T., 1990: Second Periodic Assessment of the State of the marine Environment of the Baltic Sea, 1984-1988. -Baltic Sea Environment Proceedings No. 35 B.
    Lumberg, A., Ojaveer, E., 1991: On the Environment and Zooplankton Dynamics in the Gulf of Finland in 1961-1990. Proc. Estonian Acad. Sci. Ecol. 3:131-140.
    Matthaus, W. & Schinke, H., 1994: Mean Atmospheric Circulation Patterns Associated with Major Baltic Inflows. -Deutsche Hydrogr. Zeitsch. 46:321-339.
    Ojaveer, H., Lumberg, A. & Ojaveer, E., 1998: Highlights of zooplankton dynamics in Estonian waters (Baltic Sea). - ICES J. Mar. Sci. 55:748-755.
    Ranta, E. & Vuorinen, I.; 1990: Changes of species abundance relations in marine meso-zooplankton at Seili, Northern Baltic Sea, 1967-1975. -Aqua Fennica, 20:171-180.
    Roemmich, d., McGowan, J., 1995: Climatic warming and the decline of zooplankton in the California Current. Science, 267:1324-1327
    Segerstråle, S.G., 1969: Biological fluctuations in the Baltic Sea. -Progress in Oceanography, 5:169-184.
    Viitasalo, M., Vuorinen, I. & Saesmaa, S., 1995: Mesozooplankton dynamics in the northern Baltic Sea: implications of variations in hydrography andclimate. -J.Plankton Res. 17:1857-1878.
    Vuorinen, I. Ranta, E., 1987: Dynamics of marine mesozooplankton at Seili, Northern Baltic Sea, in 1967-1975. -Ophelia 28:31-48.
    Vuorinen, I., Hänninen, J., Helminen, U., Viitasalo, M. & Kuosa, H., 1998: Proportion of copepod biomass declines with increasing salinity in the baltic Sea. -ICES J. Mar. Sci. 55:767-774.
 
[Back to Contents]

The long-term changes of plankton in the Eastern Gotland Basin (Baltic Sea) due to its eutrophication in 1976-1990

Elena  Kostrichkina, Baiba Kalveka, Anda Ikauniece
Institute of Aquatic Ecology, University of Latvia
Riga, Latvia

The eutrophication and its effect on marine biota and water quality are one of the most important contemporary problems of the Baltic Sea. The eutrophication and its consequences had specific features at various regions, determined by local nutrient loads, water exchange, hydrologic and geomorphologic properties.
    We set the analysis of one Baltic Sea region as a task for this study. The long-term dynamics of phyto- and zooplankton species composition and biomass in the Eastern Gotland Basin are described trying to find out the environmental factors responsible for these changes. As the decrease of salinity and its impact on zooplankton abundance has been reported from the other areas of the Baltic Sea (Viitasalo et al., 1995) we expected some salinity induced changes in species composition and decline in total zooplankton abundance.

Material and methods

Study area
    The Eastern Gotland Basin occupies the middle part of the Baltic Proper at the east of the Gotland. The Gotland Deep with a depth from 110-200 m and the maximal depth of 247 m is situated  in the central part of Basin. The primary thermocline and halocline divide the water masses into several layers having following specific properties (Mihailov et al., 1992a):
  1. the upper layer with the lowest salinity, under the direct influence of climatic and meteorologic factors, with drastic seasonal (<10 °C up to 15-17 °C) changes of temperature and small seasonal and interannual fluctuations of salinity (7.0 - 7.7 PSU); the size of layer in summer does not exceed 25-30 m.
  2. the cold intermediate  layer, restricted by thermocline and halocline, reflecting the condition of water after the autumn convection; water temperature is low (2-4 °C) during the year, salinity is not higher than 8.5 PSU; the size of the layer in summer is approximately 40-45 m.
  3. the deep layer under the halocline, never mixing with upper and intermediate layers, receiving the saline water inflows episodically; the water temperature is 4-6 °C, salinity - 10-13 PSU; during stagnation periods anoxic conditions arise.
In more shallow zones the deep layer is not formed. At the depth of 48 meters the annual cycle of water temperature disappears (Mihailov et al., 1992). Thus the autumn-winter convection encloses the upper 50 m layer.
    The advection of more saline North Sea waters into intermediate or deep layers increases the salinity, aerates the deep layer, changes the temperature and brings the deep pool of nutrients above the halocline.

Sampling and analysis
    Phytoplankton was sampled in the central part of the Gotland Deep at station BMP J1 in February, May, August and October-November of 1976-1990. Sampling was fulfilled with water sampler at 0.5, 10 and 20 m depth (Fig.1). Samples were analyzed under binocular microscope MBI-3, the biomass (wet weight) was calculated according to measured cell volume and the abundance of cells in 0-20 m layer.

Fig. 1 The sampling stations in the Eastern Gotland Basin

    Zooplankton was sampled at 6-8 stations in the Eastern Gotland Basin (Fig.1) at the same months as phytoplankton. Samples were gathered with Juday net (160 microns mesh size) by vertical hauls in the layers of 0-25, 25-50 and 50-100 m. The data were integrated for 0-100 m layer to avoid the impact of vertical migration, the biomass (wet weight) was calculated according to abundance and individual weights of organisms.
    Hydrological (water temperature, salinity, oxygen concentration) and hydrochemical (concentrations of phosphates, total phosphorus, nitrites and nitrates) data sampled simultaneously with plankton material at station BMP J1 were used for characterization of environment. The average values of nutrient concentrations for 0-10 m layer, of temperature and salinity for 0-20 m and 50-100 m layers, of oxygen content for 50-100 m layer and 150m depth were calculated.
    For characteristics of the solar radiation data of Riga Hydrometeorological Observatory on amount of sunshine hours in January-February at 1976-1990 were used.
    Seasonal approach was used when analyzing the long-term dynamics of  both environmental conditions and communities. The correlation method was applied for determination of effect of environmental factors.

Results

Winter
    In February of 1976-1992 the water temperature fluctuated from 0.46 to 4.10 °C in the 0-20 m layer and from 2.96 to 4.53 °C in the 50-100 m layer. The long-term dynamics of temperature in the both layers did not have any clear trends. The salinity changed from 8.15 to 7.36 PSU in the 0-20 m layer and from 10.28 to 7.75 PSU in 50-100m layer, having also strong negative trends in both layers (r2=0.69, p=0.0001, n=15 and r2=0.49, p=0.004, n=15, respectively). The oxygen concentration in 50-100 m layer in most cases exceed 3 ml/l and its long-term dynamics had a positive trend (r2=0.49, p=0.004, n=15). In deeper layer the oxygen regime regressed though. In 150 m depth since 1983 anoxic conditions were observed and negative trend (r2=0.57, p=0.001, n=15)  at this depth was stated for the whole period  under investigation.
    The long-term dynamics of salinity and oxygen concentrations had the same design in the rest of the seasons, therefore we will not describe them further on.
    Concentrations of phosphates were between 0.50 and 0.75 mmol/l and did not have common trend for the long-term dynamics. After 1984 a relative stability of values was observed. The concentrations of nitrites+nitrates were changing from 3.91 to 6.98 mmol/l. The long-term trend was not stated, some stability of values was observed after 1985. DIN/DIP ratio was considerably lower than Redfield number and fluctuated between 6.1 and 10.3, having no clear long-term tendencies.
Phytoplankton species composition in February of 1978-1987 included 31 taxons (Table 1). Diatoms, blue-green and green algae had the highest biomass. Decreasing part of diatoms and increasing significance of blue-green algae in the total biomass were the observed changes in the species structure of community. The total phytoplankton biomass fluctuated from 1.24 (in 1986) to 34.8 (in 1983) mgm-3. The relations with solar radiation were not found.
 
Season Taxonomic group Amount of species
Winter Diatomophyceae 
Dinophyceae 
Chlorophyceae 
Cyanophyceae 
Cryptophyceae
1
Spring Diatomophyceae 
Dinophyceae 
Chlorophyceae 
Cyanophyceae 
Cryptophyceae 
Euglenophyceae 
Chrysophyceae
27 
19 
1
Summer Diatomophyceae 
Dinophyceae 
Chlorophyceae 
Cyanophyceae 
Cryptophyceae 
Euglenophyceae 
Chrysophyceae
10 
15 
12 
14 
1
Autumn Diatomophyceae 
Dinophyceae 
Chlorophyceae 
Cyanophyceae 
Cryptophyceae 
Euglenophyceae
16 
10 
1
Tab. 1: The taxonomic composition of phytoplankton community in the Central Gotland Deep,  1976-1978

    Zooplankton organisms in February of 1976-1991 were represented in the main by copepods, rotifer Synchaeta baltica found in small numbers. Pseudocalanus elongatus had the highest biomass till 1981 and rapid decline of it afterwards, with strong negative trend (r2= 0.79, p=0.001, n=14) thus corresponding with the trends of salinity. The biomasses of Acartia spp. and Temora longicornis did not reach very high values but their part in the total biomass raised after Pseudocalanus decrease.The total biomass of zooplankton changed from 22 to 350 mgm-3. Same declining tendency (r2=0.81, p=0.016, n=14) after 1981 reflected the dynamics of Pseudocalanus biomass.

Spring
    In May of 1976-1992 the water temperature in 0-20 m layer changed between 3.2 °C and 8.2 °C, in 50-100 m layer - between 2.0 °C and 4.6 °C. The long-term dynamics in both layers did not had any tendencies although in 1988-1992 temperature in the upper layer was the highest during the whole period of investigations.
    The nutrient pool in May was considerably utilized already. Phosphate concentrations were between 0.06 - 0.39 mmol/l and their long-term trend was negative (r2=0.62, p=0.002, n=12). Content of nitrites+nitrates altered from 0.10 - 0.55 mmol/l but reflected no common  long-term trend. The DIN/DIP ratio was 0.5-5.0. Our data on silicate concentrations in the investigated region are only from 1985-1990. They fluctuated between 4.9-13.8 mmol/l.
    Phytoplankton species composition consisted of 61 taxons (Table 1). Diatoms and dinophlagellates were the most abundant. The considerable increase in the part of dinophlagellates (particularly - Peridiniella catenata) in the total phytoplankton biomass was the most important change in community structure. The total phytoplankton biomass showed an increasing trend (r2=0.64, p=0.0097, n=9). Long-term dynamics of phytoplankton biomass did not coincide with water temperature and nutrient concentration.
    Zooplankton species composition in spring included copepods, rotifers, Appendicularia, also cladocerans. The long-term changes in community structure manifested as drastic decrease of Pseudocalanus biomass in 1980s and significant increase in biomass of rotifers at the end of 1980s. The biomass of Acartia spp. also obtained more considerable role in total zooplankton biomass during 1980s. The total zooplankton biomass varied from 49-269 mgm-3. Its long-term tendencies had a negative trend in 1980-1987 (r2=0.60, p=0.041, n=8), coinciding with decrease of Pseudocalanus biomass, and features of increase afterwards due to high numbers of rotifers. Water temperature determined most considerably the time and intensity of zooplankton biomass development (r2=0.62, p=0.001, n=13) and salinity (r2=0.51, p=0.001, n=11) affected the level of biomass.

Summer
    In August of 1976-1992 water temperature in 0-20 m layer was between 12.6 °C and 17.5 °C, in 50-100 m layer - between 2.5 °C and 4.8 °C. The long-term dynamics in the upper layer showed no general trends, although in summers of 1983-1990 (except 1987) the values were higher than the long-term average. No trends were observed in the lower layer.
    The concentrations of phosphates were low - 0.03-0.25 mmol/l, with decrease observed in 1977 -1985 (Fig.4). Total phosphorus concentration changed from 0.33 to 0.80 mmol/l, having a tendency to increase after 1986 (Fig.4). The concentrations of nitrites+nitrates were between 0.12 and 0.45 mmol/l, without any general long-term trends.
    Phytoplankton species composition in 1976-1990 included 56 taxons (Table 1). The blue-green algae (except in 1983 and 1990) were the basic contributors to total phytoplankton biomass (Fig.2). In 1983 diatoms and in 1990 dinophlagellates were prevailing (Fig.2). The main change in community structure was the decrease of biomass of Snowella lacustris after 1981. A appearance of high abundance of Cryptophyceae after 1981 was observed although no representatives of this group occured in the formaline-fixed samples before. However, this phenomenon did not have a considerable importance in total phytoplankton biomass, which fluctuated in limits of 38.7-1234.5 mgm-3. The average biomasses in 1970s were higher than in 1980 (if not regarding 1981 and 1987).

Fig. 2 Phytoplankton biomass and species composition in August

    Zooplankton species composition in summer included more cladoceran - Bosmina longispina, Podon spp. , rotifer - Keratella spp. and meroplanktonic - Bivalvia spp. representatives, together with copepods Pseudocalanus elongatus, Acartia spp., Temora longicornis, Centropages hamatus (Fig.3). The decrease of biomass of Pseudocalanus occuring since beginning of 1980s and increase of Bosmina biomass observed in 1982-1990 were the most obvious changes in community's structure. Total zooplankton biomass fluctuated from 284 - 707 mgm-3 without a common trend. Also correspondance of biomass with long-term dynamics of temperature, salinity and food supply was not confirmed.

Fig. 3 Zooplankton biomass and species composition in August

Autumn
    In October-November the water temperature in 0-20 m layer varied from 6.7 °C to 10.9 °C, in 50-100 m layer - from 2.9 °C to 5.4 °C. The long-term tendencies were not stated as the interannual variation in both layers was high due to autumn convection.
    Considerable variation was obvious also in nutrient concentrations - 0.05 - 0.48 mmol/l for phosphates, 0.32-0.73 mmol/l for total phosphorus and 0.09 - 1.63 mmol/l for nitrites+nitrates. Clear trends were not found for these parameters, too.
Phytoplankton species composition included 48 taxons (Table 1). Diatoms, dinophlagellates, blue-green algae were forming the largest part of total biomass. A small growth in biomass of dinophlagellates in 1985-1987 and exceptional increase of Coscinodiscus granii biomass in 1988-1990 characterized the changes in the structure of community. The total phytoplankton biomass was between 36 and 1610 mgm-3 having no general long-term trend. No relationship with thermal or nutrient regimes was found.
    All abovementioned copepods, cladocerans - Evadne nordmanni, rotifers - S. baltica and Appendicularia species formed zooplankton community in autumn. The decrease of Pseudocalanus biomass was observed also in autumn since 1982. Total biomass changed from 73 to 329 mgm-3 and had a negative trend (r2= 0.73, p=0.0068, n=8) after 1982 (Fig. 6). The correspondance with salinity dynamics was found (r2=0.62, p=0.0007, n=14).

Conclusions

The increase of nutrient winter pools in the upper layer of Eastern Gotland basin stopped in the beginning of 1980s. The stableness of nutrient concentrations on relatively high levels was traced since mid-1980s. Low N/P ratios affirmed the potential nitrogen limitation.
    Most important changes in phytoplankton community structure occured at the second half of 1980s. They manifested as the decrease of diatoms and increase of dinophlagellates (in spring and summer) and blue-green algae (in summer). These changes were related with decline in spring and summer nutrient concentrations at the second half of 1980s and with increase of water temperature at the end of 1980s. The growth of Cryptophyceae abundance was observed during 1980s.
The positive trend in the total phytoplankton biomass was observed only for spring due to intensive development of dinophlagellates after 1980s. In summer of 1983-1990  as a consequence of decreased nutrient concentrations, phytoplankton biomass was lower than in 1976-1979. The increase of autumn biomass at the end of 1980s was determined  by high abundance of diatom Coscinodiscus granii.
    Salinity decrease in late 1970s caused the decline of key zooplankton species Pseudocalanus elongatus after 1981 in all seasons. The increased water temperature and favourable food conditions induced the mass occurence of Synchaeta spp. (in spring) and Bosmina longispina (in summer) at the end of 1980s.
    The long-term dynamics of total zooplankton biomass had negative trends during winter, spring and autumn after 1981. The dynamics at summer had a bimodal character: it reflected the dynamics of Pseudocalanus biomass till mid-1980s and the increase of Bosmina, Temora longicornis and Acartia spp. afterwards.
    Thus the pelagic ecosystem in spite of "stabilised" eutrophication was inconstant in the Eastern Gotland basin. It had significant quantitative and qualitative changes in planktonic communities determined by climatic conditions, salinity decrease and alterations in particular nutritional conditions of separate plankton groups.

References

    Mihailov A., Mihailov N., Volkov N., 1992a. 4.4. 4. Sezonnaja izmenchivostj [Seasonal dynamics].- Gidrometeorologija i gidrohimija morei SSSR. T.3 Baltijskoje more. Vip.1 Gidrometeorologicheskije uslovija [Hydrometeorology and hydrochemistry of the seas of the USSR. Vol.3 The Baltic Sea. No. 1 Hydrometeorological conditions]. St. Petersburg, Gidrometeoizdat, 288-310 (in Russian).
    Mihailov A., Mihailov N., Smirnova A., 1992b. 4.4.1. Obschije svedenija [General information].- Gidrometeorologija i gidrohimija morei SSSR. T.3 Baltijskoje more. Vip.1 Gidrometeorologicheskije uslovija [Hydrometeorology and hydrochemistry of the seas of the USSR. Vol.3 The Baltic Sea. No. 1 Hydrometeorological conditions]. St. Petersburg, Gidrometeoizdat, 270-271 (in Russian).
    Viitasalo M., Vuorinen I., Saesmaa S., 1995. Mesozooplankton dynamics in the northern Baltic Sea: implications of variations in hydrography and climate. - Journal of Plankton Research, 17, 1857-1878.
[Back to Contents]

The analysis of long-term data series of plankton, herring and sprat in the Central Baltic

Georgs Kornilovs
Latvian Fisheries Research Institute
Riga, Latvia

In the frames of the BASYS project (Subproject 1b: Processing of historical data) Latvian Fisheries Research Institute (LATFRI) worked on establishment of five data bases: hydrology in the Central Baltic, zooplankton in the Central Baltic, ichthyoplankton in the Central Baltic, feeding of herring and feeding of sprat. Currently the analysis of these long -term data series are performed to explain the observed variability of hydrological and biological parameters.

The oceanographic investigations in the Central part of the Baltic sea were carried out since 1960. These investigations include seasonal measurements of water temperature, salinity and oxygen content as well as meteorological observations, and cover the Eastern part of the Gotland Deep or the Latvian economical zone in the Baltic sea. In 1970s and 1980s additional more frequent (monthly) sampling was performed on some selected stations. The first analysis of long-term changes of water salinity and temperature in the Central Baltic was presented at the 2nd BASYS conference in Stockholm. Based on the seasonal observations the  oceanographic parameters were calculated for the different water layers. They characterised the main parameters in the upper layer 0 to 50 m, in the layer of halocline 70-150 m and in the deepest layer 150-200 m. Periods with an obvious trends were distinguished after analysing the material. The tendencies in the changes of salinity and temperature in the different regions and water layers coincided only partly. Two periods with the opposite tendencies were found in the upper layer: salinity was increasing until the end of the 1970s, and afterwards it was decreasing. 4 periods were found in the layer of halocline: slight increase in salinity in the 1960s, quasiconstant level of salinity in the 1970s, sharp decline in the 1980s, and rapid increase in the 1990s. In the deepest layer salinity was  decreasing until the beginning of 1990s, afterwards short-term very rapid increase of it was observed. The mean water temperature in halocline was increasing till 1977, then there was a decrease till 1988 and a rapid increase till 1991. In the 1990 the water temperature in halocline is slowly decreasing. At present the analysis is continued to seek relationships between salinity in the Central Baltic and river run-off, sea level and large-scale atmospheric circulation (climate change).

Regular investigations of zooplankton in the Central Baltic are carried out since 1960. There are four seasonal surveys every year covering a net of stations in the economical zone of Latvia (eastern part of the Gotland Deep). Until 1991 the zooplankton samples were taken also in the northern and southern parts of the Baltic sea. All zooplankton species or larger taxonomic units were determined in the samples. For Copepoda species also all stages of development (7) were determined. The abundance of certain zooplankton species depend on many factors such as water salinity, temperature, feeding conditions, predation and interaction with other species. Besides the influence of these factors is variable with seasons, years and longer periods. The hydrological environment and the predation pressure of pelagic fishes has significantly changed since the beginning of 80s. Due to the lack of strong inflows from the North sea the salinity in the Central Baltic has decreased. A row of mild winters since 1989 has changed the temperature regime especially in winter and spring. Since the end of 1980s the stock size of sprat has significanly increased due to successful reproduction in 1990s and decreased predation pressure of cod (ICES, 1998). On contrary the stock biomass of herring in the Central Baltic is gradually diminishing since the beginning of 1980s (Figure 1).

Figure 1

All these changes have their impact on the zooplankton community. In the previous studies some relationships between zooplankton and pelagic fishes have been stated. Thus the decrease of Pseudocalanus elongatus abundance was positively correlated with the decrease of mean weight-at-age of herring (Naglis and Sidrevics, 1993). In the neighbouring Gulf of Riga the abundance of Limnocalanus macrurus was negatively correlated with the stock size of the Gulf herring (Sidrevics et al, 1993). The species composition of zooplankton in the Central Baltic is rather stable while relative abundances of many species has signicantly changed during the investigation period. Ps. elongatus which was one of the dominating species till 1987 has strongly decreased in number (Figure 2).

Figure 2

On the contrary the abundance of Temora longicornis and Acartia spp. has increased still both having high interannual variability. The brackish water species Eurytemora affinis has become more frequent and abundant in the investigated area.The decrease of Ps. elongatus abundance is related to the diminishment of salinity in the Central Baltic still the influence of pelagic fish predation can not be excluded. At present long-term changes of zooplankton species abundance and their links with hydrography and other environmental factors (also large-scale atmospheric circulation) are studied using statistical downscaling.

Regular ichthyoplankton surveys (3-5 a year) are performed in the Central Baltic since 1973. Fish eggs and larvae are determined to species, counted and measured. Beginning with the mid 1980s the abundance of cod eggs and larvae in the Gotland Deep has strongly diminished. In the beginning of 1990s the eggs and larvae of cod were not found but since 1994 they are again in the samples although in very low numbers. In 1990s this region has lost its importance as spawning and reproduction area for cod. On contrary the abundance of sprat eggs and larvae has considerably increased especially in 1990s due to the increased spawning stock biomass. Other fish species are less abundant in the samples. The long term-data series are analysed with the aim to use ichthyoplankton data for prediction of year-class strength of sprat which has very high interannual variability. Besides long-term changes in the abundance and composition of ichthyoplankton in the Central Baltic will be analysed.

The feeding of pelagic fishes in the Central Baltic is studied in LATFRI since 1977. Since the beginning of 1980s the mean weight-at-age of herring started to decrease. At present in many regions of the Baltic sea the mean weight-at-age of herring is at its lowiest level since the beginning of 1970s. Since 1993 similar decrease of mean weight-at-age is observed also for sprat. Simultaneously  changes in diet of herring and sprat have been revealed. The analysis of herring feeding were presented at the 2nd BASYS conference. In this as well as in previous analysis the daily feeding activity, long-term changes of food spectrum of  herring and sprat was described (Davidyuk et al, 1992, Fetter and Davidyuk, 1996). It was concluded that during the whole period of 1977-1998 the main food components of herring Pseudocalanus elongatus and Mysis sp. were substituted by Temora longicornis. The food spectrum of sprat has not changed so drastically although in 1990s the importance of Acartia spp. has increased especially in some years while T. longicornis is still the main food item of sprat.
Significance of Ps.elongatus and T.longicornis for herring feeding during the whole period changed conversely.  In 1977-1983 Ps.elongatus was the main food for herring but after 1987 it practically vanished from herring diet. On the contrary T. longicornis being before 1982-1983 less important food item for herring, after 1985 quite strongly increased in herring diet. Also the condition factor of herring was studied in connection with herring feeding and chnges in mean-weight-at-age. Although the condition factor fluctuated it did not show so drastic decrease compared with fish weight and length. The low values of condition factor for winter were observed in 1985-1990. It was an intermediate period in the herring growth dynamics. In the period of low growth  of herring (1992-1996) the values of condition factor for winter were nearly on the  level of 1977-1983 when the growth rate of fishes was high.

At present the data of herring and sprat feeding are analysed jointly. The main goal of the work is the comparative description of feeding dynamics - sesonal and interannual- with the emphasis on the resemblance and differences between the main pelagic species. A possible food competition will be discussed according to spatial variability of food composition and feeding periodicity of herring and sprat. The estimation of daily rations and annual food consumption will be performed from the observations in the investigated period.
 

References

    Davidyuk, A., Fetter, M. and Hoziosky, S. 1992. Feeding and growth of Baltic herring (Clupea harengus m. membras L.). ICES, C.M. 1992/J:27.
    Fetter, M. and Davidyuk, A. 1993. Herring feeding and growth in the eastern Baltic during 1977-1990. ICES, C.M. 1993/J:27.
    ICES 1998. Report of the Baltic Fisheries Assessment Working Group. ICES C.M. 1998/ACFM:16.
Naglis, A. and Sidrevics, L. 1993. The analysis of mean weight-at-age of baltic herring in the eastern Baltic proper, SD 28. ICES, C.M. 1993/J:24.
    Sidrevics, L., Line, R., Berzinsh, V. and Kornilovs, G. 1993. Long-term changes of zooplankton abundance in the Gulf of Riga. ICES, C.M. 1993/L:15.
 
[Back to Contents]

On the mean circulation of the Baltic Sea

Andreas Lehmann, Institute of Marine Research Kiel, Germany
Hans­Harald Hinrichsen, Institute of Marine Research Kiel, Germany
Wolfgang Krauss, Institute of Marine Research Kiel, Germany

Introduction

It is commonly known that due to the ephemeral nature of the atmospheric conditions over the Baltic Sea current observations show high fluctuations. But this means not necessarily that there are no persistent current systems found in the Baltic Sea. From long­term current observations at different positions in the Baltic the general circulation can be derived. However, due to the enormous costs and extensive fishing activities which makes it impossible at least in some areas of the Baltic Sea to install corresponding observational systems, it is not possible to measure the general circulation. Although, surface currents have been obtained from different locations of the Baltic Sea (Sjöberg, 1992), the three­dimensional structure of the circulation is more or less unknwon. The three­dimensional circulation is not only of interest for physical ogceanographers, it is also quite important for fishery biology or biogeochemical investigations.

In order to shed some light on this problem we determined the annual averaged circulation from a coupled sea ice­ocean model run for 1992. The model has a horizontal resolution of approximately 5 km and 41 levels in the vertical which enables us to resolve the upper 100 m with levels of 3 m thickness. The model is forced by realistic atmospheric conditions taken from the SMHI meteorological data base and by river runoff taken from a monthly mean runoff data set. Prognostic variables are the 3­D current field, temperature, salinity and oxygen and the 2­D surface elvations and the barotropic tranport. These prognostic variables have been extracted from the model every 6 hours and were subsequently averaged over the whole year 1992.

General Circulation

From averaged model results (monthly or annual means) there is a clear evidence of characteristic persistent circulation patterns which comprise mostly the subbasins of the Baltic Sea with less transport between the basins (Lehmann and Hinrichsen, 1998). However, averaged currents give no information about their variability. Thus, for a more complete representation regarding the variability of currents we calculated the so called stability B. The stability of currents is defined as the ratio of the averaged vectorial velocity and the averaged arithmetic velocity.

    B = ( u2 + v2 )1/2 / ( N-1 ( Snun2 + vn2 )1/2 )
    u = N-1 Snun
    v = N-1 Snvn

with u and v the averaged components of the flow, and N is the number of current observation at the location under consideration. The vectorial mean is obtained by taking the vectorial mean value of individually observed current vectors, and the arithmetic mean velocity is obtained by averaging the speeds without regard to the current direction (Neumann and Pierson, 1967).

Fig. 1: Annual average of the barotropic circulation [cm/s] underlayed with the stability B of the barotropic flow for 1992. Colour bars show stability values 0-1.

As can be seen from Fig. 1 the most pronounced structure with the highest stability is a cyclonic circulation cell comprising the eastern Gotland Basin. Most of the water is recirculating in the eastern part, but at the northern tip there is a bifurcation into the western part. This current branch bifurcates again with one branch flowing south of  Öland into the Bornholm Basin and returning into the eastern Gotland Basin by passing through the Stolpe Trench. The second branch closes the loop into the eastern Gotland Basin through a passage between the Middle Bank and Hoburg Bank. On average the stability of the circulation pattern in the Gotland Basin is higher than 50 % which means in more than 50 % of the observations (model data
every 6 hours) this pattern is prevailing. About 1000 the main part of the water mass circulation comprising the whole Gotland Basin occur above the permanent halocline. Due to the strong current fluctuations in the Danish Straits, the stability hardly reaches values > 30%.
 

Bottom Currents

Fig. 2: Annual average of the bottom velocity [cm/s] underlayed with teh stability B of the near-bottom flow for 1992.

In Fig. 2 we present the corresponding bottom currents. The stability patterns are quite similar to those obtained for the barotropic flow. The most pronounced feature coincides with the barotropic circulation cell in the Gotland Basin with
highest stabilities due to the barotropic component of the flow. However, highest stabilities are in most cases not coupled with highest bottom velocities. It should be noted, that in those areas where the stability is high, quasi­permanent currents
will be found, whereas in those areas where the stability is low high variability of the currents occur. In areas of the deep basins where the stability and the bottom currents are small, occasionally high bottom currents can occur which may hinder
sedimentation or even lead to resuspension. However, for a resuspension of sediment, the near bottom flow must be above a critical velocity. This events may only be detected by considering the maximum bottom current velocities.
Bottom currents directed eastwards can be found in the Bornholm Gat and the Stolpe Trench with stabilities higher than 50 %. The annual average of the bottom current from the Arkona Basin into the Bornholm Basin through the Bornholm
Gat reaches values of about 10 cm/s, and through the Stolpe Trench 5 further to the east. While the compensating inflow of highly saline water through the Danish Straits is irregularly, the bottom flow through the Bornholm Gat and
Stolpe Trench appears to be a persistent signal.

Vertical Velocities

Fig. 3: Annual average of the barotropic circulation [cm/s] underlayed with the vertically averaged vertical velocity for 1992. Colour bars represent vertical velocities [w 10-4 cm/s]

From the annual average of the horizontal velocity components the vertically averaged vertical velocity can be calculated (Fig. 3). Along the western coasts of the Baltic upwelling occurs in a distance of about 20 detected. The annual average of the surface wind results in south­westerly winds more or less homogenously distributed over the Baltic Sea. The interaction of the south­westerly winds together with the basin­like bottom topography determines the areas where most likely up­ and downwelling will occur. These areas can be regarded as the most active areas in the vertical exchange of water masses of the Baltic Sea. However, some dynamical patterns like the circulation cell in the eastern Gotland Basin are coupled with diapycnal exchange although the averaged vertical velocities are not as high as along the coasts.

Conclusions

From the coupled sea ice­ocean simulations the general circulation of the Baltic Sea can be determined. In spite of the ephemeral nature of the atmospheric forcing rather stable circulation patterns occur. These circulation cells can not only be found for the simulation of 1992, the structures appear to be similar also for other years, although the strength of the circulation cells are different. From the averaged vertical velocity field, areas where most likely diapycnal mixing will occur can be identified. Near bottom velocity areas where the velocities are small and additionally the stability is high are most likely the areas where sediment deposition will occur. In those areas where the bottom velocities and the stability are small the variability of the currents is high and occasionally high bottom currents may occur which hinder sediment deposition and under certain conditions resuspension of bottom sediments can happen.

References

    Lehmann, A. and H.­H. Hinrichsen, 1998: On the thermohaline variability of the Baltic Sea. Submitted.
    Neumann, G. and W.J. Pierson, 1967: Principles of Physical Oceanography. Prentice­Hall Int. Inc., London, 545 pp.
    Sjöberg, B., 1992: Sea and Coast. National Atlas of Sweden, SMHI Norrk¨oping. SNA Publishing, 128 pp.
 
[Back to Contents]


BASYS CALENDAR
Here the BASYS-relevant meetings, workshops and conferences are notified.
 
Date Event Location Contact
15.-17.03.1999 6. MEETING SP6 (Baltic Sea Ice) Hailuoto/Icebreaker, FI alehmann@ifm.uni-kiel.de
22.-24.03.1999 6. MEETING OF SP2 (Pelagic Fluxes) IE Tallinn, EE
ulf_l@system.ecology.su.se
12.-15.04.1999 4. MEETIMG OF SP7 (Paleoenvironment) IO-PAS Sopot, PL
boris.winterhalter@gsf.fi
23.04.1999 7. MEETING OF THE SSC (Scientific Steering Committee) U Hamburg, DE bodo.bodungen@io-warnemuende.de
26.-27.04.1999 5. MEETING OF SP3A (Coastal-Basin Fluxes) U Aarhus, DK
kay.emeis@io-warnemuende.de
Spring 1999 6. MEETING OF SP4 (Nearshore and Coastline Processes) GI Greifswald, DE
kschwarzer@email.uni-kiel.de
May 1999 5. MEETING OF SP8 (System Analysis and Models) EMI Tallinn, EE
fred@sysetm.ecology.su.se
08.-09.06.1999 4. MEETING OF SP5 (Atmospheric Load) IVL Gothenburg, SE michael.schulz@dkrz.de
31.07.1999 FINAL REPORT IO Warnemünde, DE bodo.bodungen@io-warnemuende.de
01.-17.09.1999 ADVANCED STUDY COURSE: 
Marine System Analysis
IO Warnemünde, DE
karin.lochte@io-warnemuende.de,
http://www.io-warnemuende.de/Projects/Basys/events/studycourse.htm
20.-22.09.1999 3. ANNUAL BASYS CONFERENCE IO Warnemünde, DE
bodo.bodungen@io-warnemuende.de,
http://www.io-warnemuende.de/Projects/Basys/events/con3.htm
31.10.1999 OFFICIAL END OF BASYS
 
[Back to Contents]


 
PROJECT INFORMATION

BASYS - Baltic Sea System Study -

Subproject Title Coordinator Mail to
1A Coordination and Data Management Bodo von Bodungen bodo.bodungen@io-warnemuende.de
1B Processing of Historical Data Juergen Alheit juergen.alheit@io-warnemuende.de
2 Pelagic Fluxes Ulf Larsson ulf_l@system.ecology.su.se
3A Coastal-Basin Fluxes Kay-Christian Emeis kay.emeis@io-warnemuende.de
3B Basin-Basin Fluxes Matti Perttilae matti.perttilae@fimr.fi
4 Nearshore and Coastline Processes Peter Stoffers 
Klaus Schwarzer
pstoffers@email.uni-kiel.de 
kschwarzer@email.uni-kiel.de
5 Atmospheric Load Michael Schulz michael.schulz@dkrz.de
6 Baltic Sea Ice Wolfgang Krauss 
Andreas Lehmann
wkrauss@ifm.uni-kiel.de 
alehmann@ifm.uni-kiel.de
7 Paleo-Environment Boris Winterhalter boris.winterhalter@gsf.fi
8 System Analysis and Modelling Fredrik Wulff fred@system.ecology.su.se
9 Circulation and Diapycnal Exchange Gerd Becker gerd.becker@m2.hamburg.bsh.d400.de
EU Research Project
Funded by European Union RTD programmes
Marine Science and Technology MAST-3 (contract MAS3-CT96-0058) and
International Cooperation INCO (contract IC20-CT96-0080)
 
 
MAST office Project Supervision Christos Fragakis christos.fragakis@dg12.cec.be
 
[Back to Contents] 

 
IMPRESSUM
The BASYS NEWSLETTER is published 3-monthly by the BASYS secretary
(email to: christoph.zuelicke@io-warnemuende.de).
It serves for rapid communication amongst the BASYS participants, and can be noticed by the whole scientific community.
    The BASYS NEWSLETTER contains the following sections: The notes shall be very brief (about 1-2 pages, including dates, participants and contact address). The papers may be more extended (3-5 pages, figures may be included).
    For the submission of contributions HTML documents would be very welcome, but commonly used text formats could be treated as well. Figures should be supplemented extra as postscript, bitmaps of GIF files. Put the files on FTP ftp.io-warnemuende.de/pub/incoming/basys/news/ and email to: marion.sussujew@io-warnemuende.de. Each author is responsible himself for his contributions.
    Each interested person will be subscribed to an email list and will be informed via this information channel on each new issue. The NEWSLETTER is available through internet URL  http://www.io-warnemuende.de/Projects/Basys/newslett/. For further information, please contact the BASYS secretary.

(Christoph Zuelicke, January 1999, email to: christoph.zuelicke@io-warnemuende.de)
 

[Back to Contents]

last edited: 10.08.1999
Critics, ideas, suggestions to this page to:
christoph.zuelicke@io-warnemuende.de
 

Critics, ideas, suggestions to this server to:
webmaster@io-warnemuende.de